Abstract The Eastern Tropical Pacific (ETP) is characterized by complex oceanographic dynamics that affect the regional sea surface temperature (SST), a key parameter driving marine ecosystems. To date no comparative studies have been conducted on the spatial and temporal variations of SST among the several marine protected areas in the ETP. The present study used remote sensing to evaluate the monthly, annual, and decadal variations in SST from 1982 to 2019 in Coiba National Park (Coiba), Cocos Island National Park (Cocos), Malpelo Fauna and Flora Sanctuary (Malpelo), Gorgona Island National Park (Gorgona), Galápagos Marine Reserve (Galápagos) and La Plata Island (La Plata). An overall increasing trend in SST was observed across all study sites, except for West Galápagos. A decrease in SST anomaly was observed from February through April at Malpelo, Coiba, Gorgona and La Plata. These observations are likely in response to the Humboldt Current and upwelling events in the Colombian Pacific Basin. Using a longer SST dataset is recommended to determine if the annual, monthly and decadal trends observed here are driven by local, regional or global processes. Consistent monitoring of SST trends is a strategic practice in understanding and planning for current and projected changes in the marine environment.

Resumen El Pacífico Oriental Tropical (POT) se caracteriza por presentar una dinámica oceanográfica compleja que afecta la temperatura superficial del mar (TSM), un parámetro clave que influye los ecosistemas marinos. Hasta la fecha no se han realizado estudios comparativos de las variaciones espaciotemporales de la TSM entre las distintas áreas marinas protegidas en el POT. Esta investigación utilizó sensores remotos para evaluar las variaciones mensuales, anuales y decadales de la TSM desde 1982 a 2019 en el Parque Nacional Coiba (Coiba), Parque Nacional Isla del Coco (Coco), Santuario de Fauna y Flora Isla Malpelo (Malpelo), Parque Nacional Natural Isla Gorgona (Gorgona), Reserva Marina de Galápagos (Galápagos) e Isla de La Plata (La Plata). Se observó una tendencia creciente en la TSM en la mayoría de los sitios estudiados, excepto en el oeste de Galápagos. Se registró una disminución en la anomalía de la TSM durante los meses de febrero a abril en Malpelo, Coiba, Gorgona y La Plata. Las diferencias observadas son probablemente en respuesta a la corriente de Humboldt y eventos de afloramiento en la cuenca del Pacífico colombiano. Se recomienda emplear series de tiempo de mayor resolución temporal para determinar si las tendencias anuales, mensuales y decadales observadas en este estudio responden a cambios locales, regionales o globales. El monitoreo constante de las tendencias en la TSM es una práctica estratégica para comprender y planificar los cambios actuales y proyectados en el ambiente marino.

Hawksbill sea turtles (Eretmochelys imbricata) inhabiting the eastern Pacific Ocean are one of the world's most threatened marine turtle management units. Despite the fact that knowledge about the status of sea turtles at foraging grounds is a key element for developing the effective conservation strategies, comprehensive studies of hawksbills at foraging habitats in the eastern Pacific remain lacking. For many years anecdotal information indicated Coiba Island National Park in Panama as a potentially important hawksbill foraging ground, which led to the initiation of monitoring surveys in September 2014. Ongoing mark-recapture surveys to assess population status, generate demographic data and identify key foraging sites have been conducted every six months in the park since that time. To date, a total of six monitoring campaigns consisting of four days each have been conducted, leading to the capture and tagging of 186 hawksbills, 51 of which were recaptured at least once. The size range of captured individuals was 30.0 to 75.5 cm and largely comprised of juveniles. Somatic growth rates of individual hawksbills were highly variable, ranging from -0.78 to 7.1 cm year-1. To our knowledge, these are the first published growth rates for juvenile hawksbill turtles in the eastern Pacific Ocean. When these growth data are combined with information on hawksbill demography and distribution, our findings indicate Coiba Island National Park is one of the most important known foraging sites for hawksbill sea turtles in the eastern Pacific Ocean.

El Banco Hannibal se encuentra en Coiba que es Patrimonio de la Humanidad de UNESCO en el Pacífico de Panamá y es una importante zona de pesca. Este estudio evaluó las principales características topográficas y la complejidad de la montaña submarina Banco Hannibal mediante el uso de sensores remotos acústicos. A pesar del estado de protección de la zona y la importancia para las especies comerciales de peces como el pargo y el atún, la montaña submarina no ha tenido ningún estudio detallado salvo alguna colección de organismos. El área de estudio de alrededor de 125km² identificada usando cartas de navegación fue definida usando datos de sonar de barrido lateral durante julio 2008. La información batimétrica fue importada a ArcGIS donde se creó un modelo digital batimétrico DBM y mapa de pendientes. La extensión del ArcGIS Benthic Terrain Modeler BTM se utilizó para calcular el índice de posición batimétrica o BPI pendiente de cada celda relativa a las celdas vecinas y rugosidad. El banco Hannibal es un guyot alargado, triangular montaña submarina plana en parte superior, que se extiende en profundidad desde 53m hasta 416m, tiene una superficie de 76km² y es de 14.4km de largo y 7.1km de ancho. El Banco Hannibal está compuesto por pendientes pronunciadas, pendientes más suaves, parte superior de la montaña submarina, crestas altos relieves en la parte superior, zonas rugosas en las crestas, parte superior de los montes submarinos y pendiente, barrancos y crestas. El banco es asimétrico, con una pendiente relativamente suave y con surcos a través de la superficie en el lado norte en comparación con el lado suroeste que es mucho más pronunciado y más rugoso. Hay dos cumbres al norte y al sureste de la orilla, que varían en profundidad de 180 a 333m. El sustrato rocoso comprende unos 22.6km² del banco y sedimentos 37.8km². El banco y sus caras abruptas, áreas rugosas y pináculos proporcionan estructuras verticales que pueden alterar y mejorar topográficamente las corrientes. Esto se cree produce un afloramiento local que explicaría las mayores agregaciones de peces que son el objetivo de la pesca comercial y deportiva. Es probable que el Banco Hannibal tenga una presión cada vez mayor en el futuro debido al cambio climático y la pesca y este estudio ha producido información valiosa que permite la clasificación de los hábitats para el manejo de las especies asociadas y su pesca, así como la iniciación de la investigación sobre montañas submarinas de Panamá.

The Hannibal Bank sits within the Coiba UNESCO World Heritage Site in Pacific Panama and is also a fisheries management zone. Despite the protected status of the area and the importance of the Bank for commercial fish species such as snapper and tuna, the seamount has received no detailed survey except some collection of organisms. This study mapped the major topographic features and complexity of the Hannibal Bank seamount using acoustic remote sensing. A survey area of around 125km² was defined using existing charts and side-scan sonar data were collected during July 2008. A bathymetric output was imported to ArcGIS where a digital bathymetric model and slope map were created. The Benthic Terrain Modeler BTM extension for ArcGIS was used to calculate bathymetric position index and rugosity, and used to create a map of zones representing the various seabed morphology zones. The Hannibal bank is an elongated, triangular guyot flat topped seamount, which ranges in depth from 53m to 416m, covers an area of 76km² and is 14.4km long and 7.1km wide. Hannibal bank is composed of steep slopes, more gentle slopes, top of the seamount, crests elevated ridges at the top of the pinnacles, rugose areas on crests, top of seamount and slope, gullies and pinnacles. The bank is asymmetric in nature with the Northerly side having a relatively gentle slope with gullies across the surface compared to the SouthWest side which is far steeper and more rugose. There are two pinnacles to the North and South East of the bank that range in depth from 180 to 333m. Rocky substrate makes up 22.6km² of the bank and sediment 37.8km². The bank and its steeply sided, rugose areas and pinnacles provide upright structures which can disrupt and topographically enhance currents, increasing productivity. The rugose areas of Hannibal Bank should be primary targets for further research efforts as they may contain corals and their rugosity indicates that these should be some of the highest faunal diversity areas of the bank. Hannibal Bank is likely to come increasing pressure in the future through climate change and fishing and this study has produced valuable information to assist in the future mapping and management of habitats, associated species and fisheries.

Este estudo relata o primeiro registro de Molossus coibensis J. A. Allen, 1904 para a Amazônia brasileira e o segundo para o Brasil. Ele também apresenta uma análise comparativa com outras duas espécies de Molossus de pequeno porte no Brasil: M. molossus (Pallas, 1766) e M. aztecus Saussure, 1860. O estudo é baseado em oito fêmeas capturadas em áreas urbanas da cidade de Bragança, no norte do estado do Pará. Dados morfométricos e comparação qualitativa de características morfológicas permitem a identificação dos espécimes como M. coibensis, até então conhecido apenas em Tapirapuã, no sudoeste do estado de Mato Grosso, Centro-Oeste do Brasil.

This study reports on the occurrence of Molossus coibensis J. A. Allen, 1904, in the Brazilian Amazon and the second record of this species from Brazil. It also presents a comparative analysis with other two small-bodied Molossus species that occur in Brazil: M. molossus (Pallas, 1766) and M. aztecus Saussure, 1860. The report is based on 8 females captured in urban areas of the city of Bragança, in the northern Brazilian state of Pará. Morphometric data and the qualitative comparison of morphological features permited the identification of the specimens as M. coibensis, hitherto known only in Tapirapoan, in the southwestern state of Mato Grosso, west-central Brazil.

The identity of two insular populations of rodents of the nominal species, Zygodontomys brevicauda (Allen & Chapman, 1893), from the Veragua Archipelago was examined. The mitochondrial cytochrome-b gene was sequenced from specimens collected on Isla Coiba (n = 10), Isla Cébaco (n = 3) and on the nearby Peninsula Azuero (n = 3) in Panama and compared with sequences of Z. brevicauda and a number of other related species from GenBank. For Panama, phylogenetic analyses identified two clades within Zygodontomys Allen, 1897; one representing Isla Coiba and another clade composed of rats from Isla Cébaco and from the mainland on the Peninsula Azuero, as well as a GenBank sequence of Z. brevicauda from Venezuela. We suggest that the population from Isla Coiba may represent a previously undescribed species of sigmodontine rodent that is endemic to this Pacific Island.

Many studies over the past twenty years have documented the richness of arthropod galling species around the world, and some have proposed hypotheses to explain local and global patterns of galling species richness. However, few studies have been directed toward understanding how the gall-inducing species are locally distributed. The aim of this study was to determine the distribution of gall-inducing arthropods species at Coiba National Park, a tropical habitat on the Pacific coast of Panama. Our results suggest that more gall-inducing species had an aggregated distribution, and gall-inducing arthropod diversity shows a strong beta diversity component. Geographic distance was not correlated with similarity in gall-inducing species composition between the studied sites. This fact has important implications when trying to estimate gall-inducing arthropod richness and general patterns, and could cause contradictory results for hypotheses that attempt to explain the local and global patterns of galling species richness.

El interés por el estudio de las agallas y los artrópodos que las inducen ha crecido en todo el mundo en los últimos veinte años. Sin embargo, los artrópodos que inducen agallas en la región Neotropical son probablemente los menos estudiados. Un estudio de la riqueza y composición de artrópodos que inducen agallas fue desarrollado en el Parque Nacional Coiba en la Republica de Panamá. Los datos provienen de muestreos intensivos de dos semanas, efectuados entre agosto de 1997 y septiembre de 1999, en 17 sitios del área insular del Parque Nacional Coiba. Un total de 4942 agallas, que corresponden a 50 especies de insectos y nueve de ácaros, fueron colectadas en 50 especies de plantas vasculares de 30 familias. El 62.7% de las agallas correspondieron a especies de la familia Cecidomyiidae (Diptera), el 15.3% a ácaros de la familia Eriophyidae, el 8.5% a Psyllidae (Homoptera), un 6.8% a Coccidae y el 5.1% a Phlaeothripidae (Tysanoptera). Las familias de plantas con más especies de inductores de agallas fueron Myrtaceae con siete, Bignoniaceae con cinco y Euphorbiaceae, Fabaceae y Melastomataceae con cuatro. Las agallas formadas en hojas representaron el 93% del total. El número de inductores de agallas por sitio osciló entre uno y diecinueve. La riqueza de artrópodos inductores de agallas del Parque Nacional Coiba se discute con datos disponibles de la literatura para el área continental de Panamá y el Neotrópico. Nuestros resultados apoyan la idea de que es prematuro concluir que la riqueza de artrópodos que inducen agallas disminuye hacia el Ecuador.

Interest in studying galls and their arthropods inducers has been growing rapidly in the last two decades. However, the Neotropical region is probably the least studied region for gall-inducing arthropods. A study of the richness and composition of gall-inducing arthropods was carried out at Coiba National Park in the Republic of Panama. Field data come from samples obtained between August 1997 and September 1999, with three (two-week long) more intensive samplings. Seventeen sites, representing the main land habitats of Coiba National Park were surveyed. 4942 galls of 50 insect and 9 mite species inducing galls on 50 vascular plants from 30 botanical families were colleted. 62.7% of the galls were induced by gall midges (Diptera, Cecidomyiidae), 15.3% by mites, Eriophyidae, 8.5% by Homoptera, Psyllidae, 6.8% by Coccidae and 5.1% by Phlaeothripidae (Tysanoptera). The host plant families with the most galls were Myrtaceae with seven, Bignoniaceae with five and Euphorbiaceae, Fabaceae and Melastomataceae with four. Leaf galls accounted for about 93% of collected galls. Most leaf galls were pit/blister galls followed by covering and pouch galls. Gall richness per collecting site was between 1 and 19 species. Coiba’s gall diversity is discussed in relation to data available from other tropical sites from continental Panama and the Neotropical region. Our results support the idea that it may be premature to conclude that species richness of gall inducers declines near the equator. Rev. Biol. Trop. 56 (3): 1269-1286. Epub 2008 September 30.

En el Parque Nacional de Coiba, costa pacífica de Panamá, hay manglares con playas asociadas. Estudiamos la fauna de dos poliquetos orbínidos de Santa Cruz y El Gambute. En cada playa se eligieron puntos de muestreo al nivel de marea baja, media y alta y cada uno se realizaron dos muestreos, en febrero y noviembre de 1997. Encontramos individuos no identificables con certeza y una especie nueva: Orbinia oligopapillata n. sp., la cual se caracteriza por tener 15-16 setígeros torácicos con cuatro o cinco filas de uncinos y hasta tres procesos postsetales papiliformes en los lóbulos neuropodiales, por los parápodos abdominales provistos de sedas en bayoneta y por la presencia en los setígeros abdominales anteriores de papilas estomacales en escaso número. Leitoscoloplos sp. se caracteriza por llevar bolsillos laterales en los setígeros abdominales, caracter único en el género y que en la familia solo ha sido citada en dos especies del género Scoloplos. Naineris sp. se caracteriza por el número de setígeros torácicos, sus branquias, sus uncinos y sus neuropodios abdominales, bilobulados y con la acícula sobresaliente

Santa Cruz and El Gambute, two mangrove systems with associated tidal flats, were sampled in Coiba National Park, Coiba Island, Pacific of Panama. At each site, two samplings were done at low, middle and high intertidal levels in February and November of 1997. A new orbiniid species were found: Orbinia oligopapillata n. sp. is characterized by having 15-16 thoracic chaetigers with four or five rows of uncini and up to three papilliform postchaetal processes on neuropodial lobes by the abdominal parapodia bearing flail-tipped neurochaetae, and by the presence on anterior-most abdominal chaetigers of interramal cirri and a low number of subpodial and stomach papillae. A specimen belonging to genus Leitoscoloplos Day, 1977 is described as "Leitoscoloplos sp.", characterized by the lateral pouches on its abdominal chaetigers, a unique case for the family because these brooding structures have only been previously cited in two species of Scoloplos. Naineris sp. is characterized by the number of its thoracic chaetigers, branchiae, uncini and bilobed abdominal neuropodia with protruding aciculae. Rev. Biol. Trop. 54 (4): 1307-1318. Epub 2006 Dec. 15

Durante un estudio sobre la fauna marina del Parque Nacional de Coiba (Panamá) se identificaron 281 ejemplares de sílidos (Annelida: Polychaeta) intersticiales pertenecientes a 19 especies. Dos son nuevas citas para el Pacífico, Exogone (Exogone) arenosa Perkins, 1981 y Streptosyllis websteri Southern, 1914; cinco son nuevas citas para Panamá, E. (E.) dispar (Webster, 1879), E. (E.) longicornis Westheide, 1974, Salvatoria mediodentata (Westheide, 1974); Pionosyllis heterocirrata (Hartmann-Schröder, 1959) y Syllis glarearia (Westheide, 1974). Se describe un caracter no presente en la descripción original de E. Longicornis, la presencia de procesos triangulares subterminales en la primera pseudospinígera. Se describen ejemplares de la especie Syllis botosaneanui Hartmann-Schröder, 1973 con embriones en diverso estado de desarrollo en su interior, por lo que se trata de la primera referencia de esta especie como vivípara. Por último, se describe un ejemplar de Syllis sp., que se caracteriza fundamentalmente por la posesión de una larga faringe, dos lóbulos prostomiales dorsales y por sus sedas compuestas de artejos cortos y larga espinulación. Estas características diferencian a Syllis sp. de cualquier especie del género pero no se describe como especie nueva por disponerse únicamente de un solo ejemplar.

Syllids (Syllidae: Polychaeta) from Coiba National Park (Pacific of Panama). During a study carried out on soft bottoms from Coiba National Park (Panama), 218 specimens of syllids (Annelida: Polychaeta) belonging to 19 interstitial species have been identified. Two species are new reports for the Pacific Ocean, Exogone (Exogone) arenosa Perkins, 1981 and Streptosyllis websteri Southern, 1914; five are new for Panama, E. (E.) dispar (Webster, 1879); E. (E.) longicornis Westheide, 1974; Salvatoria mediodentata (Westheide, 1974); Pionosyllis heterocirrata (Hartmann-Schröder, 1959) and Syllis glarearia (Westheide, 1974). A characteristic not mentioned in the original description is herein reported for E. (E.) longicornis: the presence of triangular subterminal processes in the spiniger-like compound chaetae of chaetiger one. Specimens of the species Syllis botosaneanui Hartmann-Schröder, 1973 harboring inside embryos of various stages of development have been found; this is the first report for this species as viviparous. Lastly, one specimen of the genus Syllis has been found that is mainly characterized by its long pharynx, two dorsal prostomial lobes, and compound chaetae with short blades and long spinulation. Because we consider that one specimen is not sufficient to describe a new species we refer it as Syllis sp. Rev. Biol. Trop. 54 (3): 725-743. Epub 2006 Sept. 29.

Dentro del programa para el inventario de la fauna y flora del Parque Nacional de la Isla de Coiba (Panamá), se presenta un primer listado de los ácaros oribátidos (Acari: Oribatei) que habitan en el parque. El material objeto de este estudio, procede de diversos muestreos edáficos, incluyendo hojarasca y humus, realizados en el sitio en julio de 1998. Para un total de 1 053 ejemplares colectados en las muestras se han identificado 134 especies diferentes, de las cuales, practicamente todas son nuevas para Panamá y 8 especies se citan por vez primera en la región neotropical. Se da un nuevo nombre: Aeroppia mariehammerae n.nom. para Aeroppia sp. (Hammer, 1962) y una combinación nueva para Protoribates antillensis (Mahunka, 1985) n. comb. (Xylobates)

Taking part of the Program for The Inventory of Flora and Fauna of Coiba Island National Park (Panamá), a first list of oribatid mites (Acari: Oribatei) is presented. Materials studied come from several soil samplings, including humus and fallen leaves, done in Coiba Island in July 1998. 134 species were identified from a total amount of 1 053 individuals collected. Almost all the identified species are new records for Panamá as well as eigth species that represent first records for the Neotropical Region. A new name is given: Aeroppia mariehammerae n. nom. for Aeroppia sp. and a new combination is proposed: Protoribates antillensis (Mahunka, 1985) n. comb. (Xylobates)

During the years 1996 and 1997, a research project aiming to characterize the marine benthic fauna of Coiba National Park, located off the Pacific coast of Panamá, was conducted and special attention was paid to two environments: sandy sublittoral bottoms and tidal flats. Sublittoral samples were taken in September of 1996, from two sites located near coral reefs: Mali Rock and Granito de Oro. In both cases, sediment was composed of coarse sand coming from coral destruction. Tidal samples were collected from Santa Cruz and El Gambute, two flats associated with mangrove systems where upper, middle and lower intertidal sampling sites were selected. Three samplings were carried out in the flats, in June of 1996, in February of 1997 and in November of 1997. The study of the Paraonidae (Polychaeta) fauna revealed the presence of nine species belonging to three different genera, four of which were new records for the Pacific area encompassed between southern Mexico and northern Colombia. Moreover, a new species of Aricidea Webster, 1879 is described; it can be distinguished from the already described species within the genus by the combined presence of an extremely long antenna, which extends to chaetiger 9, of a pair of large posterior pair eyes, of 16 pairs of branchiae, and of numerous pseudoarticulated chaetae on abdominal neuropodia. These chaetae have a thick shaft with a slightly curved tip and bear a straight and very long spine inserted subterminally.

Durante los años 1996 y 1997 se realizó un proyecto de investigación destinado a caracterizar la fauna marina del Parque Nacional de Coiba, situado en la costa pacífica de Panamá, y en el que se prestó una atención especial a dos ambientes: los fondos arenosos sublitorales y la zona intermareal de las playas. Las muestras sublitorales se tomaron en septiembre de 1996, en dos localidades próximas a arrecifes de coral: Mali Rock y Granito de Oro. En ambos casos el sedimento estaba constituido por arena gruesa proveniente de la destrucción del coral. Las muestras intermareales se colectaron en Santa Cruz y El Gambute, dos playas asociadas a manglares en las que se seleccionaron puntos en el intermareal superior, medio e inferior. En las playas se realizaron tres muestreos, en junio de 1996, en febrero de 1997 y en noviembre de 1997. El estudio de la fauna de la familia Paraonidae (Polychaeta) reveló la presencia de nueve especies pertenecientes a tres géneros, cuatro de las cuales fueron nuevas citas para la región del Pacífico oriental comprendida entre el sur de México y norte de Colombia. Además, se describe una nueva especie de Aricidea Webster, 1879, que se caracteriza por poseer en combinación una antena extremadamente larga, que puede alcanzar el setígero 9, por el gran tamaño de los ojos del par posterior, por presentar 16 pares de branquias y por la presencia en los neuropodios abdominales de numerosas sedas pseudoarticuladas, que presentan un manubrio grueso y con la punta apenas curvada y una espina recta y de gran longitud insertada subterminalmente.

En cuatro campañas científicas al Parque Nacional de Coiba, Panamá, (1996-1998), se han hallado 19 especies de la familia Syllidae (subfamilias Autolytinae, Eusyllinae y Exogoninae). Se citan por primera vez para el Pacífico oriental cuatro especies: Eusyllis lamelligera Marion & Bobretzky, 1875, Exogone exmouthensis Hartmann-Schröder, 1980, Proceraea cf. cornuta (Agassiz, 1863) y Sphaerosyllis (S.) magnidentata Perkins, 1981. Diez especies se citan por primera vez para las costas del Pacífico de América Central: Amblyosyllis cf. granosa Ehlers, 1897 (sensu Westheide 1974), Amblyosyllis speciosa Izuka, 1912, Autolytus multidenticulatus Westheide, 1974, Exogone naidinoides Westheide, 1974, Grubeosyllis concinna Westheide, 1974, Odontosyllis fulgurans, (Audouin & Milne Edwars, 1833), Opisthodonta mitcheli Kudenov & Harris, 1995, Pionosyllis articulata Kudenov & Harris, 1995, Sphaerosyllis (S.) californiensis Hartman, 1966, Syllides cf. reishi Dorsey, 1978. La especie más dominante en las muestras de Pocillopora ssp. muerto, es S. armillaris con más del 50% de los ejemplares identificados y en las muestras de Telesto cf. multiflora las especies más abundante es G. concinna (67.36%). Por otro lado, una cuarta parte de las especies que aparecen en las muestras tomadas están restringidas a las aguas cálidas del Pacífico. Un 22% de las especies aparecen en Atlántico y Pacífico, y más del 13% son aparentemente circuntropicales y otro tanto aparentemente cosmopolitas.

This is the second paper on the Syllidae (Polychaeta) from hard substrata of the Coiba National Park (Pacific coast of Panama) (the first paper treated Syllinae). On this second paper nineteen species belonging to eleven genera of the subfamilies Autolytinae, Eusyllinae and Exogoninae (Syllidae: Polychaeta) are reported. The samples were collected by SCUBA diving, either by removing 4 kg blocks of dead coral (Pocillopora spp.) or scraping off 25 x 25 cm quadrats of Telesto multiflora or algae (Dyctiota cf. flavellata, Padina cf. durvillaei and another currently unidentified species). They were collected during four expeditions carried out between 1996 and 1998. Four species are newly reported for the Eastern Pacific: Eusyllis lamelligera Marion & Bobretzky, 1875, Exogone exmouthensis Hartmann-Schröder, 1980, Proceraea cf. cornuta (Agassiz, 1863) and Sphaerosyllis (S.) magnidentata Perkins, 1981. Ten species are reported for the first time from the Central America Pacific coast: Amblyosyllis cf. granosa Ehlers, 1897 (sensu Westheide 1974), Amblyosyllis speciosa Izuka, 1912, Autolytus multidenticulatus Westheide, 1974, Exogone naidinoides Westheide, 1974, Grubeosyllis concinna (Westheide, 1974), Odontosyllis fulgurans (Audouin & Milne Edward, 1833), Opisthodonta mitchelli Kudenov & Harris, 1995, Pionosyllis articulata Kudenov & Harris, 1995, Sphaerosyllis (S.) californiensis Hartman, 1966 and Syllides cf. reishi Dorsey, 1978. The family Syllidae is represented in the Coiba National Park by 14% of cosmopolitan species, 14% of circumtropical species, 22% of Atlantic and Pacific species and 25% of Pacific species restricted to warm waters. In samples of dead coral the dominant species are S. armillaris (Muller, 1771) and S. gracilis (Campoy, 1982); in samples of telestaceans Grubeosyllis concinna dominates.

Los anélidos poliquetos del Pacífico panameño en general y los sílidos en particular son poco conocidos. En el marco de un inventario faunístico del Parque Nacional de Coiba, entre 1996 y 1998 el grupo del Laboratorio de Biología Marina de la Universidad Autónoma de Madrid realizó cuatro campañas científicas en las que se ha estudiado la poliquetofauna de los fondos duros. Las muestras corresponden a 4 kg de coral muerto (Pocillopora ssp.), raspados de 25 x 25 cm de Telesto multiflora y 25 x 25 cm de algas (Dyctiota cf. flavellata, Padina cf. durvillaei y otras), siempre tomadas por buceadores con escafandra autónoma. Se presentan en este trabajo los resultados del estudio de los poliquetos de la subfamilia Syllinae Grube, 1850 (Syllidae: Polychaeta). Se describe una nueva especie, Syllis castroviejoi. Se citan por primera vez para el Pacífico oriental cuatro especies: Syllis beneliahuae (Campoy & Alquezar, 1982), S. botosaneanui (Hartmann-Schröder, 1973), S. corallicola Verrill, 1900 y S. garciai (Campoy, 1982). Dos especies se citan por primera vez para las costas del Pacífico de América Central: Branchiosyllis pacifica Rioja, 1941 and Syllis truncata Haswell, 1920. Dos especies, Syllis magna (Westheide, 1974) and S. pigmentata (Chamberlin, 1919) son nuevas para Panamá.

Four expeditions were made to Coiba National Park between 1996 and 1998 to characterize this part of the poorly known Panamian Pacific polychaete fauna. The samples were collected by SCUBA diving, either by removing 4 kg blocks of dead coral (Pocillopora sp.) or scraping off 25 x 25 cm quadrats of Telesto multiflora or algae (Dyctiota cf. flavellata, Padina cf. durvillaei and another currently unidentified species). This paper deals with the Syllinae from hard substrates. Twenty-two species belonging to six genera of the subfamily Syllinae Grube, 1850 (Syllidae: Polychaeta) are reported. A new species, Syllis castroviejoi, is described. Four species are newly reported for the Eastern Pacific: Syllis beneliahuae (Campoy & Alquézar, 1982), S. botosaneanui (Hartmann-Schröder, 1973), S. corallicola Verrill, 1900 and S. garciai (Campoy, 1982), and, likewise, two species are first reports for the Central American Pacific coast: Branchiosyllis pacifica Rioja, 1941 and Syllis truncata Haswell, 1920. Two species, Syllis magna (Westheide, 1974) and S. pigmentata (Chamberlin, 1919), are newly reported for Panama.