Abstract Lagostomus maximus is a native rodent of South America that presents economic and biological importance. However, few studies on parasites of this rodent are available throughout its geographic distribution. The aim of this study was to explore, describe and compare the structure of helminth communities from 2 populations of L. maximus in semi-captive (SCHP) and wild (WHP) conditions in Buenos Aires Province, Argentina. The structure of helminth communities was studied considering ecological data at different levels. Seven helminth taxa were collected. Graphidioides spp. and Lagostonema ecasiense were the most prevalent species in SCHP, and Viannella viscaciae in WHP. This last species showed the highest values of mean intensity and mean abundance in both host populations. Helminths from SCHP showed higher values of abundance, mean intensity, mean abundance, diversity, and evenness, and lower values of specific richness and dominance than WHP. Specific richness, evenness and dominance allowed separating the 2 host populations. The abundances of V. viscaciae and Graphidioides spp. distinguished both host populations, suggesting possible influences of human intervention and/or environmental characteristics, as consequence of semi-captive and wild conditions.

Resumen Lagostomus maximus es un roedor autóctono de América del Sur que presenta importancia económica y biológica. Sin embargo, se dispone de pocos estudios sobre sus parásitos en toda su distribución geográfica. El objetivo de este estudio fue explorar, describir y comparar la estructura de las comunidades de helmintos de 2 poblaciones de L. maximus en la Provincia de Buenos Aires, Argentina, en condiciones de semicautividad (SCHP) y silvestría (WHP). La estructura de las comunidades de helmintos se estudió considerando datos ecológicos a diferentes niveles. Se colectaron 7 taxones de helmintos. Graphidioides spp. y Lagostonema ecasiense fueron las especies más prevalentes en SCHP y Viannella viscaciae en WHP. Esta última especie mostró los valores más altos de intensidad media y abundancia media en ambas poblaciones hospedadoras. Los helmintos de SCHP mostraron valores más altos de abundancia, intensidad media, abundancia media, diversidad y equitatividad, y valores más bajos de riqueza específica y dominancia que WHP. La riqueza específica, la equitatividad y la dominancia permitieron separar a las 2 poblaciones hospedadoras. Las abundancias de V. viscaciae y Graphidioides spp. distinguieron a ambas poblaciones hospedadoras, sugiriendo posibles influencias de la intervención humana y/o características ambientales, como consecuencia de las condiciones de semicautividad y silvestría.

Abstract The similarity between parasites communities’ decay with distance and its analysis may explain important ecological process such host dispersion. Patagonia is inhabited by two armadillo species, Chaetophractus villosus and Zaedyus pichiy. In this study we describe and analyze the variation on helminth fauna of these armadillos in Patagonia compared with northern localities described in previous studies. A total of 49 armadillos were collected in Patagonia. Quantitative descriptors of parasite ecology were calculated and community structure of helminths was analyzed following the central-satellite species hypothesis. The parasite richness in Patagonia decreases almost 50% in both armadillos. Zaedyus pichiy present the same central species in Patagonia as in northern localities. For C. villosus central-satellite species analysis could not be applied. The loss of some helminths in Z. pichiy could be the result of lower temperatures or the absence of intermediate arthropods hosts. But in C. villosus the absence of some helminths with Patagonian distribution could be explained by its recent dispersion in Patagonia. Trichohelix tuberculata still being the only helminth in C. villosus introduced population of Tierra del Fuego.

Resumo Os Xenarthra, endêmicos das Américas, consistem em dois clados distintos, Pilosa (tamanduás e preguiças) e Cingulata (tatus). A associação entre xenartros e humanos remonta aos tempos antigos. Embora essas associações não pareçam ter impactado negativamente todas as populações selvagens, a caça provavelmente contribuiu para a extinção de alguns xenartros nos tempos arqueológico-geológicos e, mais recentemente, para o esgotamento ou extinção das populações locais. Sem dúvida, as atividades humanas estão impactando cada vez mais as populações selvagens de muitas espécies de xenartros. De acordo com a última avaliação da Lista Vermelha de Espécies Ameaçadas da IUCN, 15 das 30 espécies de Xenarthra são classificadas como Menos Preocupantes (LC), 5 como Deficientes em Dados (DD), 5 como Quase Ameaçadas (NT) e 5 são incluídas em uma categoria ameaçada, com uma (Bradypus pygmaeus) sendo considerado Criticamente em Perigo (CR) e quatro (Myrmeco- phaga tridactyla, Bradypus torquatus, Priodontes maximus . Tolypeutes tricinctus) designados como Vulneráveis (VU). Aqui, discutimos os desafios taxonômicos atuais do trabalho com xenartros e os impactos desses desafios na conservação desses animais. Avaliamos as principais ameaças que afetam os Xenarthra agora e no futuro, identificamos algumas iniciativas de conservação bem-sucedidas e destacamos a importância das iniciativas de especialistas individuais. Nossos objetivos são resumir o status atual dos xenartros e facilitar o desenvolvimento e a implementação de estratégias de conservação para esses animais.

Abstract The Xenarthra, which are endemic to the Americas, consist of two distinct clades, the Pilosa (anteaters and sloths) and the Cingulata (armadillos). The association between xenarthrans and humans dates back to ancient times. Although these associations do not appear to have negatively impacted all wild populations, hunting probably contributed to the extinction of some xenarthrans in archaeological-geological times and, more recently, to the depletion or extinction of local populations. Undoubtedly, human activities are increasingly impacting the wild populations of many xenarthran species. According to the last assessment for the IUCN Red List of Threatened Species, 15 of the 30 species of Xenarthra are classified as Least Concern, five as Data Deficient, five as Near Threatened, and five are included in a threatened category, with one (Bradypus pygmaeus) being considered Critically Endangered, and four (Myrmecophaga tridactyla, Bradypus torquatus, Priodontes maximus, and Tolypeutes tricinctus) designated as Vulnerable. Here, we discuss the current taxonomic challenges of working with xenarthrans and the impacts of those challenges on the conservation of these animals. We evaluate the main threats affecting the Xenarthra now and in the future, we identify some successful conservation initiatives, and we highlight the importance of initiatives by individual experts. Our goals are to summarize the current status of xenarthrans and to facilitate the development and implementation of conservation strategies for these animals.

Armadillos are among the most common mammals in agroecosystems in Argentina. Their insectivorous/omnivorous food habits raise the question about their putative role as pest controllers. The aim of this study is to describe the prey items of five armadillo species and evaluate their possible role as natural pest controllers. The stomach contents of 12 Dasypus hybridus, 10 Chaetophractus vellerosus, 14 Chaetophractus villosus, 4 Tolypeutes matacus and 9 Zaedyus pichiy were analyzed. We described the diet and identified prey items to family level, whenever possible, and computed the frequency of occurrence (FO) and relative abundance of prey items. With these values, the Item Categorization Index (ICI) was calculated to classify the items in order of importance (primary, secondary and tertiary items). Shannon diversity index was also computed. The armadillos studied here consumed arthropod species belonging to families of phytosanitary importance such as Acrididae, Scarabaeidae, Tenebrionidae, Lepidoptera, Formicidae, and Termitidae. Only two armadillos showed primary items in their diet; Z. pichiy (ants and tenebrionids) and T. matacus (termites and scarabid larvae). Important pest arthropods had FO values close to or greater than 50% in the diet of all armadillos. Armadillos consumed arthropods that are important pests for agriculture, forestry, and wood construction. Even though this study was mainly focused on armadillos’ diet, we suggest that the presence of these native species could benefit the productivity and health of agroecosystems by reducing the need for harmful agrochemicals.

We present a revised checklist of the mammals of Argentina, incorporating recent taxonomical changes and new records of mammals for the country produced since the publication of a previous list in 2006. We recordered 409 native species, corresponding to 181 genera, 46 families and 12 orders, plus 23 introduced taxa with wild populations.

Se presenta una lista revisada de los mamíferos de Argentina, incorporando los cambios taxonómicos recientes y los nuevos registros para el país producidos desde la publicación de un listado previo en 2006. Se registraron 409 especies nativas, correspondientes a 181 géneros, 46 familias y 12 órdenes, más 23 especies introducidas con poblaciones silvestres.

The long-nosed armadillos of the Dasypus genus are the richest and more widespread extant xenarthra. In 1933, the prominent Argentinean mammalogist José Yepes studied specimens collected by Salvador Mazza and named a new species: D. mazzai. Several authors interpreted that the holotype of the species was a juvenile of D. novemcinctus. In 1995, Vizcaíno support the synonymy with D. novemcinctus but accepted the existence of an endemic species of northwestern Argentina that dedicated to Yepes (D. yepesi). Morphological studies have questioned the synonymy of the species with D. novemcinctus. In this contribution we reviewed the identity of the type material by sequencing a DNA fragment of 212 bp, product of the 16S ribosomal unit of Dasypodidae. Both samples of the holotype (bone and muscle) and of a specimen from the north of Santa Fe province have a 100% similarity with the D. yepesi / D. sabanicola sequences available in the GenBank. These results indicate that the type specimen of D. mazzai is neither a D. novemcinctus nor another species of the Dasypus genus previously named. Dasypus mazzai is revalidated and the eastern boundary in the Chaco region is confirmed. Field work is required to confirm the conservation status and current distribution of this endemic mulita.

Las mulitas del género Dasypus representan el género más rico y ampliamente distribuido de los xenartros actuales. En 1933 el destacado mastozoólogo José Yepes estudió los especímenes colectados por Salvador Mazza y nominó una nueva especie: D. mazzai. Varios autores interpretaron que el holotipo era un ejemplar juvenil de D. novemcinctus. En 1995, Vizcaíno aceptó la sinonimia con D. novemcinctus, pero reconoció la existencia de una especie endémica del noroeste de Argentina que dedicó a Yepes (D. yepesi). Estudios morfológicos han cuestionado la sinonimia de esta especie con D. novemcinctus. En esta contribución se revisó la identidad del material tipo secuenciando un fragmento de ADN de 212 bp, que codifica parte de la unidad ribosomal 16S de Dasypodidae. Ambas muestras del holotipo (hueso y músculo) y de un espécimen del norte de la provincia de Santa Fe presentan 100% de similitud con secuencias de D. yepesi/D. sabanicola disponibles en el GenBank. Los resultados indican que el ejemplar tipo de D. mazzai no pertenece a D. novemcinctus ni a otra de las especies del género Dasypus previamente nominadas. Se revalida D. mazzai y se confirma el límite oriental de su distribución en la región chaqueña. Se requieren trabajos de campo para confirmar el estado de conservación y el rango de distribución actual de la especie.

ABSTRACT The aim of this work was to obtain the first estimates of survival rates (S), capture probability (p) and life expectancy for armadillos in South America by analyzing capture-mark-recapture data obtained from a population of Chaetophractus vellerosus (Gray, 1865) located in Magdalena, Buenos Aires, Argentina. From June 2006 to June 2011, we conducted 16 field surveys that resulted in 365 capture events of 152 adult C. vellerosus. For the survival analysis we used a Cormack-Jolly-Seber (CJS) modelling framework. Interannual variation in S made an important contribution to overall variation in the survival rate of C. vellerosus. The average life expectancy for females and males after attaining sexual maturity was estimated at 1.70 and 1.65 years respectively. The period of lowest survival probability was associated with dry seasons that might have affected the availability of food. This study provides the first estimates of demographic parameters for xenarthrans in South America.

During the last decades, the Pampas Region of Argentina has witnessed an expansion of the croplands at the expense of grasslands and pastures. As a result of no-till system, after harvesting the region shows large areas with stubbles. The aim of this study is to describe the assemblage of land birds and compare species richness and abundance at the community guild levels in crop and livestock fields during two contrasting periods of the year, when croplands remains at stubbles stage. Twenty-five sites were selected and in each one four samplings were conducted: two in spring-summer and two in autumn. In each site and each sampling, two crop and two livestock fields were chosen, totalling 392 fields. Birds were recorded in each field using a linear transect of 700 m by 100 m. Seventy-five species were recorded. Livestock fields supported greater species richness than crop fields throughout the study, provided by highest insectivore species richness. In spring-summer, there was a higher abundance in livestock fields, mainly of insectivore birds. There were not significant differences in species richness and abundance of granivorous birds between land uses, even though in autumn there was a sharp differential use of crop fields by the granivorous species Zenaida auriculata, which accounted for 36% of total abundance in crop fields during that period. The results show that there is an impoverishment of the bird assemblage in crop fields with regard to livestock fields and it can be expected that these trends will be accentuated as long as the substitution between these different land uses deepens.

En las últimas décadas, los agroecosistemas de la Región Pampeana en Argentina han sido objeto de cambios en los patrones de uso de la tierra caracterizados por la expansión de la agricultura en detrimento de pastizales y pasturas. Como resultado de la amplia adopción de la práctica de siembra directa, luego de la cosecha se registran en algunas zonas extensas áreas cubiertas con rastrojos. El objetivo de este trabajo es describir el ensamble de aves terrestres y comparar la riqueza específica y la abundancia a nivel comunitario y gremial de lotes agrícolas y ganaderos durante dos períodos contrastantes del año en los cuales los lotes agrícolas se encuentran en rastrojo. Se seleccionaron 25 sitios que fueron visitados en cuatro oportunidades: dos veces en primavera-verano y dos veces en otoño. En cada sitio y muestreo se eligieron dos lotes agrícolas y dos ganaderos independientes (392 lotes en total). En cada lote se registraron las aves en una transecta lineal de 700 m de largo y 100 m de ancho. Se registró un total de 75 especies. Los lotes ganaderos tuvieron una riqueza específica mayor que los agrícolas a lo largo de todo el estudio, aportada por una mayor riqueza de aves insectívoras. En primavera-verano hubo una mayor abundancia de aves en lotes ganaderos, particularmente de aves insectívoras. No hubo diferencias en la riqueza y la abundancia de aves granívoras entre lotes, aunque en el otoño se registró un marcado uso diferencial de los lotes agrícolas por parte de la especie granívora Zenaida auriculata, que acumuló el 36% de la abundancia en esos lotes agrícolas durante ese período. Los resultados muestran un empobrecimiento del ensamble de aves en lotes agrícolas con respecto a los ganaderos; es de esperar que estas tendencias se acentúen en la medida que se profundice la sustitución entre estos usos de la tierra.

We update the distribution of three species of armadillos (Chaetophractus villosus, C. vellerosus and Dasypus hybridus) in the Pampas region within the Province of Buenos Aires, Argentina, with information that comes from fieldworks conducted during 2011-2013. Armadillos were recorded through direct observations and indirect evidence (burrows and foraging holes). We obtained 632 records (10 C. vellerosus, 184 D. hybridus, and 438 C. villosus), which provide information of 27 counties for which there were no documented records of the species, we correct previous data, and we complete information of areas scarcely surveyed in previous years such as the center and western areas of the province.

Actualizamos la distribución de tres especies de armadillos (Chaetophractus villosus, C. vellerosus y Dasypus hybridus) presentes en la región pampeana comprendida en la provincia de Buenos Aires, Argentina, con muestreos de campo realizados durante 2011-2013. Los armadillos fueron registrados mediante observaciones directas y evidencias indirectas (cuevas y hozaduras). Se obtuvieron 632 registros (10 C. vellerosus, 184 D. hybridus y 438 C. villosus), lo que aporta información de 27 partidos donde no había registros documentados de estas especies, corregimos datos previos y completamos información del centro y oeste de la provincia, áreas que habían sido poco muestreadas anteriormente.

Patagonia, south of 40° S, is currently inhabited by Chaetophractus villosus and Zaedyus pichiy. In order to reconstruct the Late Pleistocene-Holocene history of these armadillos in southern South America, we compiled and discussed data from zooarcheological sites, notes from naturalists and travellers, museum specimens, and field data. We found that both species experienced significant range expansion southwards. Based on the obtained evidence we concluded that: (a) Southern Patagonia was mainly free of armadillos during the Late Pleistocene-Early Holocene; (b) in the last 10 ky Z. pichiy progressively invaded southern Patagonia from the north, down to the Santa Cruz River (~50° S); at the same time, C. villosus colonized the northern portion of Patagonia north of Chubut River (~44° S); (c) during the last century, Z. pichiy surpassed the Santa Cruz River barrier while C. villosus colonized all Patagonian territories to reach the Magellan Strait and finally was introduced in Tierra del Fuego. Although these armadillos have metabolic, dietary and behavioral adaptations to cold and dry habitats that possibly helped them to extend its range southwards, we propose that the observed distributional changes of the last century were triggered mainly by anthropogenic causes.

Patagonia, al sur de los 40° S, está actualmente habitada por Chaetophractus villosus y Zaedyus pichiy. Con el objeto de reconstruir la historia de estos armadillos en la porción austral de América del Sur durante el Pleistoceno tardío-Holoceno, compilamos y discutimos datos de registros zooarqueológicos, observaciones de naturalistas y viajeros, especímenes de museos y datos de campo. Encontramos que ambas especies experimentaron una considerable expansión de su distribución hacia el sur. De acuerdo a la evidencia obtenida concluimos que: (a) Patagonia austral estuvo básicamente libre de armadillos durante el Pleistoceno tardío- Holoceno temprano; (b) durante los últimos 10 ka, Z. pichiy invadió progresivamente el sur de la Patagonia desde el norte alcanzando el río Santa Cruz (~50° S); al mismo tiempo C. villosus colonizó la porción norte de Patagonia llegando al norte del río Chubut (~44° S); (c) durante el siglo pasado Z. pichiy superó la barrera impuesta por el río Santa Cruz mientras que C. villosus colonizó todo el territorio patagónico hasta alcanzar el estrecho de Magallanes y finalmente fue introducido en Tierra del Fuego. Aunque estas especies de armadillos tienen adaptaciones metabólicas, de dieta y de comportamiento a los ambientes fríos y secos que posiblemente les ayudaron a extender su área de distribución hacia el sur, proponemos que los cambios distribucionales de la última centuria fueron gatillados principalmente por causas antrópicas.

The home range of the screaming hairy armadillo Chaetophractus vellerosus was estimated and its spatial pattern described under different degrees of land use pressure. Fieldwork was performed between 2006 and 2011 on a cattle ranch in Magdalena (Buenos Aires , Argentina), in two paddocks of similar size but with different intensities of use (North: low; South: high). A total of 213 individuals were captured (102 males and 111 females). Data from 30 adult individuals (12 males and 18 females) were analyzed in order to determine the home range using the minimum convex polygon method. The spatial pattern was described based on 85 captures and the nearest neighbor method. No significant differences in home range size were observed between sexes and seasons (fall-winter: males 0.23 ha, females 0.27 ha; spring-summer: males 0.75 ha, females 0.13 ha). The spatial distribution of armadillos was aggregated in both paddocks in fall-winter, being higher in the paddock with low intensity of use. An aggregated distribution was also observed during spring-summer in the southern paddock, while a random arrangement was found in the northern paddock. The home range of C. vellerosus is not affected by sex or season. The spatial arrangement of individuals is influenced by the intensity of land use.

Se estimó el área de acción del armadillo Chaetophractus vellerosus y se describió su patrón espacial teniendo en cuenta la presión de uso del ambiente. Los muestreos se realizaron en un campo ganadero de Magdalena (Buenos Aires, Argentina), entre 2006 y 2011, en dos potreros de tamaño similar pero con distintas intensidades de uso (norte: baja; sur: alta). Se efectuaron 517 capturas de 213 individuos (102 machos y 111 hembras). Para determinar el área de acción se analizaron datos de 30 individuos adultos (12 machos y 18 hembras) mediante el método del mínimo polígono convexo. El patrón espacial se describió con 85 capturas y el método del vecino más cercano. No existieron diferencias significativas en el área de acción promedio entre sexos, ni entre estaciones (otoño-invierno: machos 0.23 ha, hembras 0.27 ha; primavera-verano: machos 0.75 ha, hembras 0.13 ha). Los individuos tuvieron una disposición agregada para ambos potreros en otoño-invierno, siendo mayor en el potrero de baja intensidad de uso. Durante primavera-verano en el potrero sur mantuvieron la misma distribución agregada, mientras que en el potrero norte presentaron una disposición al azar. El área de acción de C. vellerosus no se ve afectada por el sexo y las estaciones. La distribución espacial de los individuos es influenciada por la intensidad de uso del ambiente.

The aim of this study was to analyze the parasitic fauna of a population of Chaetophractus vellerosus from Magdalena, Buenos Aires province, Argentina, by parasitological analysis. We evaluated the influence on the parasitic dynamics of different host parameters -age, sex, height and weight- as well as seasonality, vegetation and human impact. Oocysts of 2 species of Eimeria (Protozoa-Coccidea) and eggs of Aspidodera sp. Trichostrongylina, Ancylostoma sp. and Trichuris sp. (Nematoda) were found. Eimeria sp. and Aspidodera sp. were the most prevalent parasites, and Ancylostoma sp. was the lowest prevalent. Parasite loads of coccideans were higher in hosts young. Parasite loads of Trichuris sp. were higher in males. Only Trichostrongylina had greater loads in specimens with poor body condition. Trichostrongylina and Eimeria sp. were most prevalent in summer, while Aspidodera sp. was in winter. Neither vegetation nor human impact influenced in parasitic loads. This study enabled us to know the parasitic fauna of a population of C. vellerosus (Dasypodidae) in Argentina through coproparasitological analysis, and monitor it for 4 consecutive years.

El objetivo de este trabajo fue analizar la fauna parasitaria de una población de Chaetophractus vellerosus de Magdalena, provincia de Buenos Aires, Argentina, mediante análisis coproparasitológicos. Se evaluó la influencia de distintos parámetros del hospedero -edad, sexo, talla y peso- estacionalidad, cobertura vegetal y el impacto antrópico sobre la dinámica de la estructura parasitaria. Se hallaron ooquistes de 2 especies de Eimeria (Protozoa-Coccidea) y huevos de Aspidodera sp., Trichostrongylina, Ancylostoma sp. y Trichuris sp. (Nematoda). Los parásitos de mayor prevalencia fueron Eimeria sp. y Aspidodera sp., el de menor, Ancylostoma sp. Las cargas parasitarias de los coccidios fueron mayores en los hospederos juveniles; las de Trichuris sp. fueron mayores en los machos. Sólo los Trichostrongylina tuvieron mayores cargas en los hospederos con mala condición corporal. Trichostrongylina y Eimeria sp. fueron más prevalentes en verano, mientras que Aspidodera sp. lo fue en invierno. Ni la cobertura vegetal ni el impacto antrópico influenciaron sobre las cargas parasitarias. El presente estudio permitió conocer la fauna parasitaria de una población de C. vellerosus (Dasypodidae) en Argentina mediante el análisis coproparasitológico y su seguimiento durante 4 años consecutivos.

This contribution presents new data about the natural history of the Dasypus hybridus (Desmarest, 1804) (Mammalia, Xenarthra, Dasypodidae). A seasonal monitoring using the capture and release method was performed over the course of three years in a 100 ha of four cattle farms in the Buenos Aires province, Argentina. Data were collected on food habits, space and time use, behavior, thermoregulation, population data, and morphology. A total of 71 captures were made. The main food items recorded were plant matter, ants and coleopterans, and did not occur seasonal differences in food habits. Dasypus hybridus concentred their activities during daytime, and were recorded a low frequency of activities during cold seasons (fall and winter). The long-nosed armadillo prefer humid soils and grasslands with high vegetation cover, and selected forest to refuge. The southern long-nosed armadillo were asocial. Rectal temperature was positively correlated with environmental temperature. The sex proportion was near to one, and no sexual dimorphism was observed. In general, the results of this study are in agreement with previous observations of other species of the same genera, but is necessary to highlight the trend observed in food habits and thermoregulatory strategy. This work provides new information about different aspects of a poor studied species that occur in a habitat subjected to high pressure to modification and use.

En este trabajo se presentan nuevos aportes sobre la historia natural de la mulita pampeana Dasypus hybridus (Desmarest, 1804) (Mammalia, Xenarthra, Dasypodidae). Los estudios de campo fueron llevados a cabo en 100 ha de cuatro establecimientos agropecuarios de la provincia de Buenos Aires, Argentina. Durante tres años se realizó un muestreo estacional de armadillos por captura y liberación. Se obtuvieron datos de hábitos alimentarios, uso del espacio y del tiempo, comportamiento, termorregulación, datos poblacionales y morfológicos. Se realizaron 71 capturas. En la dieta el ítem principal registrado fue material vegetal, seguido por hormigas e insectos coleópteros; no se observó una diferencia estacional en los hábitos alimentarios. La actividad de las mulitas se concentra durante el día, existió una baja en la frecuencia de observación durante las estaciones frías (otoño e invierno). La mulita pampeana prefiere suelos húmicos, terrenos altos y pastizales densos y altos; asimismo seleccionan los montes para refugiarse. Son individuos asociales. La temperatura rectal mostró correlaciones positivas con la temperatura ambiente. La proporción de sexos fue cercana a uno y no se observó dimorfismo sexual. Los resultados obtenidos concuerdan parcialmente con lo observado para otras especies del género, destacando las tendencias observadas en los hábitos alimentarios y en la estrategia termorregulatoria. Este trabajo representa un aporte en varios aspectos de una especie poco estudiada en una zona bajo importantes presiones de uso y modificación de hábitat.