We describe a new species of Cyrtoneuropsis Malloch, 1925 (Diptera, Muscidae) from Brazil. The new species is included in a previously published key to species. Additionally, we review a previously published matrix to Cyrtoneurina Giglio-Tos, 1893 and Cyrtoneuropsis. A total of 52 character states were modified and the new species was added to it. The modified matrix was analyzed under equal and implied weights. In all topologies, Cyrtoneuropsis spiloptera (Wiedemann, 1830) came up as the sister group of the new species. The monophyly of Cyrtoneurina and Cyrtoneuropsis were recovered, but the relationships among their species, in our data, differ from those obtained in the past.

In a study on Fanniidae biogeography, Dominguez & Roig-Juñent (2011) argued that the family had a Pangeic origin, Late Jurassic/early Cretaceous (~146 Ma). However, recent literature on Diptera supports that Schizophora radiation occurred during Cenozoic. Fanniidae is a widespread taxon and it was interpreted under the maximum vicariance paradigm; the consequence was an analysis with no alternative hypothesis, but Pangeic origin. We verified that Fanniidae historical narrative was incongruent with the Gondwana sequential break-up. A second analysis, assuming the Fanniidae origin during early Paleocene (65 Ma), showed congruence with recent geological events and with the Muscidae diversification, a closely related Muscoidea family. Our hypothesis suggests that the Fanniidae originated in Paleogene and they were affected by few events of vicariance and several expansions during Cenozoic.

En un estudio sobre biogeografía de Fanniidae, Domínguez & Roig-Juñent (2011) argumentaron que la familia era de origen Pangeico, Jurásico superior/Cretáceo inferior (~146 Ma). Sin embargo, literatura reciente sobre Diptera, confirma que la radiación de Schizophora ocurrió durante el Cenozoico. Fanniidae es un taxón ampliamente distribuido y fue interpretado bajo el paradigma de máxima vicarianza; la consecuencia, fue un análisis sin hipótesis alternativas, pero de origen pangeico. Nosotros verificamos que la narrativa histórica de Fanniidae es incongruente con la quiebra secuencial de Gondwana. Un segundo análisis, asumiendo el origen de Fanniidae durante el Paleoceno inferior (65 Ma), mostró congruencia con eventos geológicos recientes y con la diversificación de Muscidae, una familia de Muscoidea próximamente relacionada. Nuestra hipótesis sugiere que Fanniidae se originó en el Paleógeno y fueron afectados por pocos eventos de vicarianza y muchas expansiones durante el Cenozoico.